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Email replies default to Business Hub account

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We have several mailboxes set up on our instance with identically configured mailbox roles for each one.  in the last couple of weeks we've discovered that if we have the mailbox role Mailbox - BusinessHub on our list of user account roles, whenever we reply to an email it will default to that email address as the sending address irrespective of what mailbox it was sent to.


Eg.  Email sent to Service Desk. 

I reply to the email and it shows the sending account as being Business Hub instead of Service Desk.

We can manually change this of course, but surely we shouldn't have to.

Please advise why this is happening.  It looks like there is a default email setting somewhere that automatically chooses a set account to reply with rather than using the account the email was sent to.

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